动物模型与人类重大疾病机理重点实验室知识库

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	A Normalization-Free and Nonparametric Method Sharpens Large-Scale Transcriptome Analysis and Reveals Common Gene Alteration Patterns in Cancers
	Li QG1,3; He YH1,3; Li Y5; Sun C6; Wang XT7; Zhou JM2; Li HP8; Chen YB*2; Wu H1,3,9; Kong QP*1,3; Yang CP2; Pu SY1,3; kongqp@mail.kiz.ac.cn; ybchen@mail.kiz.ac.cn
	2017
发表期刊	Theranostics
卷号	7期号:11页码:2888-2899
合作性质	其它
摘要	Heterogeneity in transcriptional data hampers the identification of differentially expressed genes (DEGs) and understanding of cancer, essentially because current methods rely on cross-sample normalization and/or distribution assumption—both sensitive to heterogeneous values. Here, we developed a new method, Cross-Value Association Analysis (CVAA), which overcomes the limitation and is more robust to heterogeneous data than the other methods. Applying CVAA to a more complex pan-cancer dataset containing 5,540 transcriptomes discovered numerous new DEGs and many previously rarely explored pathways/processes; some of them were validated, both in vitro and in vivo, to be crucial in tumorigenesis, e.g., alcohol metabolism (ADH1B), chromosome remodeling (NCAPH) and complement system (Adipsin). Together, we present a sharper tool to navigate large-scale expression data and gain new mechanistic insights into tumorigenesis.
关键词	Cross-value Association Analysis Normalization-free Pan-cancer Transcriptome Heterogeneity
资助者	This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He).
收录类别	其他
语种	英语
资助者	This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He). ; This work was supported by the Key Research Program of the Chinese Academy of Sciences (Grant NO. KJZD-EW-L14), grants from the Chinese Academy of Sciences (QYZDB-SSW-SMC020), the National Natural Science Foundation of China (81272309, 81322030, U1502224, 81672764, 31271579, 81602346, 81671404, 81500670), Yunnan Applied Basic Research Projects (2014FA038), and grant from the Youth Innovation Promotion Association of Chinese Academy of Sciences (to Y.H. He).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/11821
专题	科研部门_动物模型与人类重大疾病机理重点实验室 遗传资源与进化国家重点实验室 科研部门_分子人类学（孔庆鹏） 科研部门_肿瘤信号转导（陈勇彬） 科研部门_基因调控与表达遗传（周巨民）
通讯作者	kongqp@mail.kiz.ac.cn; ybchen@mail.kiz.ac.cn
作者单位	1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China 2.Key Laboratory of Animal Models and Human Disease Mechanisms, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China 3.KIZ/CUHK Joint Laboratory of Bioresources and Molecular Research in Common Diseases, Kunming 650223, China 4.Department of Pathology, the Second Xiangya Hospital, Central South University, Changsha 410013, China 5.Farm Animal Genetic Resources Exploration and Innovation Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu 611130, China 6.Laboratory for Conservation and Utilization of Bio-Resources, Yunnan University, Kunming 650091, China 7.School of Life Sciences, University of Science and Technology of China, Hefei 230027, China 8.Key Laboratory of Computational Biology, CAS-MPG Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai 200031, China 9.Kunming College of Life Science, University of Chinese Academy of Sciences, Beijing 100049, China.
推荐引用方式 GB/T 7714	Li QG,He YH,Li Y,et al. A Normalization-Free and Nonparametric Method Sharpens Large-Scale Transcriptome Analysis and Reveals Common Gene Alteration Patterns in Cancers[J]. Theranostics,2017,7(11):2888-2899.
APA	Li QG.,He YH.,Li Y.,Sun C.,Wang XT.,...&ybchen@mail.kiz.ac.cn.(2017).A Normalization-Free and Nonparametric Method Sharpens Large-Scale Transcriptome Analysis and Reveals Common Gene Alteration Patterns in Cancers.Theranostics,7(11),2888-2899.
MLA	Li QG,et al."A Normalization-Free and Nonparametric Method Sharpens Large-Scale Transcriptome Analysis and Reveals Common Gene Alteration Patterns in Cancers".Theranostics 7.11(2017):2888-2899.

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2017071101A.pdf（1985KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文

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